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Ophioglossum polyphyllum

A.Braun ex Seub. (1844)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Ophioglossales Family: Ophioglossaceae Genus: Ophioglossum

adder's tongue, pololei

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Indigenous
Kaho'olawe Unknown
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Bibliography

Name Published In: Fl. Azor.: 17 (1844)

Other References

Palmer 2003:199 (K, O, Mo, L, M, Ka, H); Cooke Jr. 1268a BISH (Mo†, 1924); Stemmermann 3715B BISH (L†, 1979); Note: no BISH spms. from Ka, as of Mar 2025, record retained (Maya LeGrande pers. comm.)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Ophioglossum polyphyllum Kau, Hilea-Kai, radius of 10' on heiau ~30' from ocean, full sun, dry PRESERVED_SPECIMEN Greenwell, A.B.H. Degener 21241 Hawaii BISH 3/9/1950
2 Ophioglossum polyphyllum ditch in the center of the oahu country club dried ditch PRESERVED_SPECIMEN d.f. grether 4234 WIS 2/22/1946
3 Ophioglossum polyphyllum Kamakaipo sand dunes PRESERVED_SPECIMEN Cooke Jr., C.M. 1268a Molokai BISH 1/1/1924
4 Ophioglossum polyphyllum Ninole, Wailua, Kau gravelly spot on basalt point, by old Hawaiian house site PRESERVED_SPECIMEN St. John, H. 23952 Hawaii BISH 12/25/1949
5 Ophioglossum polyphyllum puako In partly shaded lawn area mowed weekly. PRESERVED_SPECIMEN funk, e.j. collector number: s.n. Oahu BISH 12/18/1992
6 Ophioglossum polyphyllum kamilo, ka`u PRESERVED_SPECIMEN Hawaii US
7 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2748a Kauai PTBG 2/5/1988
8 Ophioglossum polyphyllum Manele and Hulopoe Bays, between sand dunes, kiawe forest PRESERVED_SPECIMEN Stemmermann, R.L. 3715B Lanai BISH 2/18/1979
9 Ophioglossum polyphyllum barking sands, between kohole point and base housing Dodonaea shrubland on low, coastal sand dunes, in full sun or in thatch PRESERVED_SPECIMEN wilson, k.a. gustafson, r.; flynn, t. collector number: 1612 Kauai BISH 3/6/1989
10 Ophioglossum polyphyllum kahue, puna district In moist black soil, in sink hole 2m deep with Sida. Semi arid pahoehoe slopes. PRESERVED_SPECIMEN st.john, h. collector number: 25466 Hawaii BISH 12/24/1954
11 Ophioglossum polyphyllum manele and hulopoe bays, between sand dunes, kiawe forest PRESERVED_SPECIMEN stemmermann, r.l. herbst, d.r.; palmer, d.; corn, c. collector number: 3715b Lanai BISH 2/18/1979
12 Ophioglossum polyphyllum Kauai PRESERVED_SPECIMEN Brodie, A. s.n. Kauai BISH
13 Ophioglossum polyphyllum east maui, s coast, hanamanioa point fine coral and lava rubble PRESERVED_SPECIMEN hobdy, r.w. collector number: 2459 Maui BISH 12/1/1985
14 Ophioglossum polyphyllum Kau, Keauhou back of high tide mark PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 10/1/1931
15 Ophioglossum polyphyllum barking sands, between kohole point and base housing Dodonaea shrubland on low, coastal sand dunes, under canopy PRESERVED_SPECIMEN flynn, t. lorence, d.h.; crum, e.; momohara, d. collector number: 2683 Kauai BISH 1/15/1988
16 Ophioglossum polyphyllum Barking Sands, between Kohole Point and base housing Dodonaea shrubland on low, coastal sand dunes, under canopy PRESERVED_SPECIMEN Flynn, T. 2683 Kauai BISH 1/15/1988
17 Ophioglossum polyphyllum W Maui, Wailuku Sand hills. PRESERVED_SPECIMEN s.n. Maui BISH
18 Ophioglossum polyphyllum near sisal PRESERVED_SPECIMEN cooke jr., c.m. collector number: 1763.o Oahu BISH 2/16/1912
19 Ophioglossum polyphyllum Kahue, Puna District In moist black soil, in sink hole 2m deep with Sida. Semi arid pahoehoe slopes. PRESERVED_SPECIMEN St. John, H. 25466 Hawaii BISH 12/24/1954
20 Ophioglossum polyphyllum behind cove W of Kanaio beach hot cinders PRESERVED_SPECIMEN Hobdy, R.W. 414 Maui BISH 12/27/1978
21 Ophioglossum polyphyllum hulopoe sand under Prosopis PRESERVED_SPECIMEN herbst, d.r. collector number: 6312 Lanai BISH 2/18/1979
22 Ophioglossum polyphyllum Kahue, Puna moist black soil, sink hole 2 m deep with Sida, semiarid pahoehoe slopes PRESERVED_SPECIMEN St. John, H. 25466 Hawaii BISH 12/24/1954
23 Ophioglossum polyphyllum behind cove w of kanaio beach hot cinders PRESERVED_SPECIMEN hobdy, r.w. collector number: 414 Maui BISH 12/27/1978
24 Ophioglossum polyphyllum ruins 2 miles S of Pohie Bay soil on pahoehoe PRESERVED_SPECIMEN Higashino, P.K. 10207 Hawaii BISH 3/26/1984
25 Ophioglossum polyphyllum near Sisal PRESERVED_SPECIMEN Cooke Jr., C.M. 1763.O Oahu BISH 2/16/1912
26 Ophioglossum polyphyllum oahu PRESERVED_SPECIMEN remy, m.j. collector number: 58 Oahu BISH 1851-01-01
27 Ophioglossum polyphyllum Hulopoe sand under Prosopis PRESERVED_SPECIMEN Herbst, D.R. 6312 Lanai BISH 2/18/1979
28 Ophioglossum polyphyllum unknown PreservedSpecimen Judd, J.R. Collector Number: s.n. BISH Specimen
29 Ophioglossum polyphyllum ninole, wailua, kau gravelly spot on basalt point, by old Hawaiian house site PRESERVED_SPECIMEN st.john, h. hatheway, w.h.; morton, n.e. collector number: 23952 Hawaii BISH 12/25/1949
30 Ophioglossum polyphyllum Puako Under buffelgrass [Cenchrus ciliaris L.] PRESERVED_SPECIMEN Funk, E.J. s.n. Hawaii BISH 12/18/1992
31 Ophioglossum polyphyllum East Maui, Wailuku District, Keokea, southeast of Puu O Kali. Vegetation: Erythrina, Diospyros, Sophora, Myoporum, Euphorbia, Sideroxylon, Polyscias, Nothocestrum, Sida, Chenopodium, Nototrichium, Dodonaea, Capparis, Cocculus, Sicyos, Pellaea, Doryopteris, Ipomoea, Argemone, Waltheria, Prosopis, Melia, Leucaena, Neonotonia, Melinis, Verbesina, Lantana. PRESERVED_SPECIMEN Oppenheimer, H.L. H111320 Maui BISH 11/25/2013
32 Ophioglossum polyphyllum w maui, wailuku Sand hills. PRESERVED_SPECIMEN bailey, e. collector number: s.n. Maui BISH
33 Ophioglossum polyphyllum unknown PreservedSpecimen Munro, G.C. Collector Number: 167 BISH Specimen 4/2/1905
34 Ophioglossum polyphyllum unknown PreservedSpecimen Kanashiro, T. Collector Number: 5730 BISH Specimen 12/11/1947
35 Ophioglossum polyphyllum kau, hilea-kai, radius of 10' on heiau ~30' from ocean, full sun, dry PRESERVED_SPECIMEN greenwell, a.b.h. collector number: degener 21241 Hawaii BISH 3/9/1950
36 Ophioglossum polyphyllum East Maui, S coast, Hanamanioa Point fine coral and lava rubble PRESERVED_SPECIMEN Hobdy, R.W. 2459 Maui BISH 12/1/1985
37 Ophioglossum polyphyllum ruins 2 miles s of pohie bay PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10207 BISH 3/26/1984
38 Ophioglossum polyphyllum kamakaipo sand dunes PRESERVED_SPECIMEN cooke jr., c.m. collector number: 1268a Molokai BISH 1/1/1924
39 Ophioglossum polyphyllum hawaiian islands. PRESERVED_SPECIMEN d. d. baldwin s.n. GH
40 Ophioglossum polyphyllum kauai Exposed slopes, mostly bare ground with sparse scattering of wind-trimmed Schinus, Metrosideros & Styphelia PRESERVED_SPECIMEN brodie, a. Oahu BISH
41 Ophioglossum polyphyllum Barking Sands, between Kohole Point and base housing Dodonaea shrubland on low, coastal sand dunes, in full sun or in thatch PRESERVED_SPECIMEN Wilson, K.A. 1612 Kauai BISH 3/6/1989
42 Ophioglossum polyphyllum w maui, wailuku sand hills PRESERVED_SPECIMEN Maui BISH
43 Ophioglossum polyphyllum south kohala district. collected east of queen kaahumanu highway between mauna lani drive and puako turn off Growing in shady forest on steep, talus slope near valley bottom. Associated vegetation: Aleurites moluccana, Psidium guajava, Pouteria sandwicensis, Freycinetia arborea, Antidesma platyphyllum, Pipturus albidus, Cyrtandra grayi, Cyrtandra munroi, Tectaria gaudichaudii, Strongylodon ruber, Christella parasitica, Blechnum appendiculatum. PRESERVED_SPECIMEN funk, e.j. collector number: s.n. Maui BISH 2/4/1991
44 Ophioglossum polyphyllum W Maui, Wailuku Sand hills. PRESERVED_SPECIMEN Bailey, E. s.n. Maui BISH
45 Ophioglossum polyphyllum unknown PRESERVED_SPECIMEN judd, j.r. collector number: s.n. Oahu BISH
46 Ophioglossum polyphyllum Oahu PRESERVED_SPECIMEN Remy, M.J. 58 Oahu BISH 1/24/1905
47 Ophioglossum polyphyllum kau, keauhou PRESERVED_SPECIMEN judd jr., a.f. BISH 10/1/1931
48 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2749 Kauai PTBG 2/5/1988
49 Ophioglossum polyphyllum unknown Lowland Dry Shrubland at 1800 ft on a W facing slope with Dodonaea viscosa, Sida fallax, Lipochaeta lavarum, L. lobata and Eragrostis sp. PRESERVED_SPECIMEN munro, g.c. collector number: 167 Maui BISH
50 Ophioglossum polyphyllum Kamoku. PRESERVED_SPECIMEN Munro, G.C. 167 Lanai BISH 4/2/1905
51 Ophioglossum polyphyllum Wailuku PRESERVED_SPECIMEN Bailey, E. s.n. Maui BISH 1879-00-00
52 Ophioglossum polyphyllum PRESERVED_SPECIMEN d. d. baldwin 113 MU
53 Ophioglossum polyphyllum Polihale State Park, dunes in front of reef zone near Queens Pond. Coastal dry shrubland mixed community, sand dunes and lithified sand outcrops. PRESERVED_SPECIMEN K. R. Wood 18381 Kauai PTBG 2/11/2020
54 Ophioglossum polyphyllum N. Kona; upper makai slopes of Puu Oo, Land of Maniniowale, mauka of Queen Kaahumanu Hwy Ca. 100 individuals near summit, in exposed sites in grassland of Pennisetum setacum. PRESERVED_SPECIMEN Nagata, K.M. 4270 Hawaii BISH 11/22/1993
55 Ophioglossum polyphyllum unknown PRESERVED_SPECIMEN Judd, J.R. s.n. BISH 1888-00-00
56 Ophioglossum polyphyllum n. kona; upper makai slopes of puu oo, land of maniniowale, mauka of queen kaahumanu hwy PRESERVED_SPECIMEN nagata, k.m. collector number: 4270 Kauai BISH 11/22/1993
57 Ophioglossum polyphyllum Kanaio, near Piimoe Hill, above Pohakea; slopes all the way to Hwy have scattered plants, eastern side of Kanaio. Elev. 1170-1460 ft. Alien vegetation. PRESERVED_SPECIMEN Steve Perlman 13309 Maui PTBG 2/4/1993
58 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2683 Kauai PTBG 1/15/1988
59 Ophioglossum polyphyllum Polihale State Park. Dune system behind beach. Low shrubland. On open dunes or in sand pockets of lithified dunes. PRESERVED_SPECIMEN Tim Flynn 3285 Kauai PTBG 3/7/1989
60 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2698 Kauai PTBG 1/18/1988
61 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2697 Kauai PTBG 1/18/1988
62 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2706 Kauai PTBG 1/18/1988
63 Ophioglossum polyphyllum Polihale State Park, dunes in front of reef zone near Queens Pond. Coastal dry shrubland mixed community, sand dunes and lithified sand outcrops. PRESERVED_SPECIMEN K. R. Wood 15850 Kauai PTBG 2/27/2014
64 Ophioglossum polyphyllum U. S. Navy Pacific Missile Range Facility at Barking Sands. PRESERVED_SPECIMEN Tim Flynn 5185 Kauai PTBG 1/11/1993
65 Ophioglossum polyphyllum East Maui, Keokea, SE of Puu O Kali. Lava flow. PRESERVED_SPECIMEN Hank Oppenheimer H111320 Maui PTBG 11/25/2013
66 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2700 Kauai PTBG 1/18/1988
67 Ophioglossum polyphyllum US Navy Pacific Missle Range Facility. Barking Sands. Just north of Nohili Point; base of sand dunes 200 ft inland from beach. Open areas with Leucaena-Prosopis shrubland. PRESERVED_SPECIMEN Walter Odening s.n. Kauai PTBG 1/31/1990
68 Ophioglossum polyphyllum Kau, Hilea-Kai, radius of 10' on heiau ~30' from ocean, full sun, dry PRESERVED_SPECIMEN Greenwell, A.B.H. Degener 21241 Hawaii BISH 3/9/1950
69 Ophioglossum polyphyllum ditch in the center of the oahu country club dried ditch PRESERVED_SPECIMEN d.f. grether 4234 WIS 2/22/1946
70 Ophioglossum polyphyllum Kamakaipo sand dunes PRESERVED_SPECIMEN Cooke Jr., C.M. 1268a Molokai BISH 1/1/1924
71 Ophioglossum polyphyllum Ninole, Wailua, Kau gravelly spot on basalt point, by old Hawaiian house site PRESERVED_SPECIMEN St. John, H. 23952 Hawaii BISH 12/25/1949
72 Ophioglossum polyphyllum puako In partly shaded lawn area mowed weekly. PRESERVED_SPECIMEN funk, e.j. collector number: s.n. Oahu BISH 12/18/1992
73 Ophioglossum polyphyllum kamilo, ka`u PRESERVED_SPECIMEN Hawaii US
74 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2748a Kauai PTBG 2/5/1988
75 Ophioglossum polyphyllum Manele and Hulopoe Bays, between sand dunes, kiawe forest PRESERVED_SPECIMEN Stemmermann, R.L. 3715B Lanai BISH 2/18/1979
76 Ophioglossum polyphyllum barking sands, between kohole point and base housing Dodonaea shrubland on low, coastal sand dunes, in full sun or in thatch PRESERVED_SPECIMEN wilson, k.a. gustafson, r.; flynn, t. collector number: 1612 Kauai BISH 3/6/1989
77 Ophioglossum polyphyllum kahue, puna district In moist black soil, in sink hole 2m deep with Sida. Semi arid pahoehoe slopes. PRESERVED_SPECIMEN st.john, h. collector number: 25466 Hawaii BISH 12/24/1954
78 Ophioglossum polyphyllum manele and hulopoe bays, between sand dunes, kiawe forest PRESERVED_SPECIMEN stemmermann, r.l. herbst, d.r.; palmer, d.; corn, c. collector number: 3715b Lanai BISH 2/18/1979
79 Ophioglossum polyphyllum Kauai PRESERVED_SPECIMEN Brodie, A. s.n. Kauai BISH
80 Ophioglossum polyphyllum east maui, s coast, hanamanioa point fine coral and lava rubble PRESERVED_SPECIMEN hobdy, r.w. collector number: 2459 Maui BISH 12/1/1985
81 Ophioglossum polyphyllum Kau, Keauhou back of high tide mark PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 10/1/1931
82 Ophioglossum polyphyllum barking sands, between kohole point and base housing Dodonaea shrubland on low, coastal sand dunes, under canopy PRESERVED_SPECIMEN flynn, t. lorence, d.h.; crum, e.; momohara, d. collector number: 2683 Kauai BISH 1/15/1988
83 Ophioglossum polyphyllum Barking Sands, between Kohole Point and base housing Dodonaea shrubland on low, coastal sand dunes, under canopy PRESERVED_SPECIMEN Flynn, T. 2683 Kauai BISH 1/15/1988
84 Ophioglossum polyphyllum W Maui, Wailuku Sand hills. PRESERVED_SPECIMEN s.n. Maui BISH
85 Ophioglossum polyphyllum near sisal PRESERVED_SPECIMEN cooke jr., c.m. collector number: 1763.o Oahu BISH 2/16/1912
86 Ophioglossum polyphyllum Kahue, Puna District In moist black soil, in sink hole 2m deep with Sida. Semi arid pahoehoe slopes. PRESERVED_SPECIMEN St. John, H. 25466 Hawaii BISH 12/24/1954
87 Ophioglossum polyphyllum behind cove W of Kanaio beach hot cinders PRESERVED_SPECIMEN Hobdy, R.W. 414 Maui BISH 12/27/1978
88 Ophioglossum polyphyllum hulopoe sand under Prosopis PRESERVED_SPECIMEN herbst, d.r. collector number: 6312 Lanai BISH 2/18/1979
89 Ophioglossum polyphyllum Kahue, Puna moist black soil, sink hole 2 m deep with Sida, semiarid pahoehoe slopes PRESERVED_SPECIMEN St. John, H. 25466 Hawaii BISH 12/24/1954
90 Ophioglossum polyphyllum behind cove w of kanaio beach hot cinders PRESERVED_SPECIMEN hobdy, r.w. collector number: 414 Maui BISH 12/27/1978
91 Ophioglossum polyphyllum ruins 2 miles S of Pohie Bay soil on pahoehoe PRESERVED_SPECIMEN Higashino, P.K. 10207 Hawaii BISH 3/26/1984
92 Ophioglossum polyphyllum near Sisal PRESERVED_SPECIMEN Cooke Jr., C.M. 1763.O Oahu BISH 2/16/1912
93 Ophioglossum polyphyllum oahu PRESERVED_SPECIMEN remy, m.j. collector number: 58 Oahu BISH 1851-01-01
94 Ophioglossum polyphyllum Hulopoe sand under Prosopis PRESERVED_SPECIMEN Herbst, D.R. 6312 Lanai BISH 2/18/1979
95 Ophioglossum polyphyllum unknown PreservedSpecimen Judd, J.R. Collector Number: s.n. BISH Specimen
96 Ophioglossum polyphyllum ninole, wailua, kau gravelly spot on basalt point, by old Hawaiian house site PRESERVED_SPECIMEN st.john, h. hatheway, w.h.; morton, n.e. collector number: 23952 Hawaii BISH 12/25/1949
97 Ophioglossum polyphyllum Puako Under buffelgrass [Cenchrus ciliaris L.] PRESERVED_SPECIMEN Funk, E.J. s.n. Hawaii BISH 12/18/1992
98 Ophioglossum polyphyllum East Maui, Wailuku District, Keokea, southeast of Puu O Kali. Vegetation: Erythrina, Diospyros, Sophora, Myoporum, Euphorbia, Sideroxylon, Polyscias, Nothocestrum, Sida, Chenopodium, Nototrichium, Dodonaea, Capparis, Cocculus, Sicyos, Pellaea, Doryopteris, Ipomoea, Argemone, Waltheria, Prosopis, Melia, Leucaena, Neonotonia, Melinis, Verbesina, Lantana. PRESERVED_SPECIMEN Oppenheimer, H.L. H111320 Maui BISH 11/25/2013
99 Ophioglossum polyphyllum w maui, wailuku Sand hills. PRESERVED_SPECIMEN bailey, e. collector number: s.n. Maui BISH
100 Ophioglossum polyphyllum unknown PreservedSpecimen Munro, G.C. Collector Number: 167 BISH Specimen 4/2/1905
101 Ophioglossum polyphyllum unknown PreservedSpecimen Kanashiro, T. Collector Number: 5730 BISH Specimen 12/11/1947
102 Ophioglossum polyphyllum kau, hilea-kai, radius of 10' on heiau ~30' from ocean, full sun, dry PRESERVED_SPECIMEN greenwell, a.b.h. collector number: degener 21241 Hawaii BISH 3/9/1950
103 Ophioglossum polyphyllum East Maui, S coast, Hanamanioa Point fine coral and lava rubble PRESERVED_SPECIMEN Hobdy, R.W. 2459 Maui BISH 12/1/1985
104 Ophioglossum polyphyllum ruins 2 miles s of pohie bay PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10207 BISH 3/26/1984
105 Ophioglossum polyphyllum kamakaipo sand dunes PRESERVED_SPECIMEN cooke jr., c.m. collector number: 1268a Molokai BISH 1/1/1924
106 Ophioglossum polyphyllum hawaiian islands. PRESERVED_SPECIMEN d. d. baldwin s.n. GH
107 Ophioglossum polyphyllum kauai Exposed slopes, mostly bare ground with sparse scattering of wind-trimmed Schinus, Metrosideros & Styphelia PRESERVED_SPECIMEN brodie, a. Oahu BISH
108 Ophioglossum polyphyllum Barking Sands, between Kohole Point and base housing Dodonaea shrubland on low, coastal sand dunes, in full sun or in thatch PRESERVED_SPECIMEN Wilson, K.A. 1612 Kauai BISH 3/6/1989
109 Ophioglossum polyphyllum w maui, wailuku sand hills PRESERVED_SPECIMEN Maui BISH
110 Ophioglossum polyphyllum south kohala district. collected east of queen kaahumanu highway between mauna lani drive and puako turn off Growing in shady forest on steep, talus slope near valley bottom. Associated vegetation: Aleurites moluccana, Psidium guajava, Pouteria sandwicensis, Freycinetia arborea, Antidesma platyphyllum, Pipturus albidus, Cyrtandra grayi, Cyrtandra munroi, Tectaria gaudichaudii, Strongylodon ruber, Christella parasitica, Blechnum appendiculatum. PRESERVED_SPECIMEN funk, e.j. collector number: s.n. Maui BISH 2/4/1991
111 Ophioglossum polyphyllum W Maui, Wailuku Sand hills. PRESERVED_SPECIMEN Bailey, E. s.n. Maui BISH
112 Ophioglossum polyphyllum unknown PRESERVED_SPECIMEN judd, j.r. collector number: s.n. Oahu BISH
113 Ophioglossum polyphyllum Oahu PRESERVED_SPECIMEN Remy, M.J. 58 Oahu BISH 1/24/1905
114 Ophioglossum polyphyllum kau, keauhou PRESERVED_SPECIMEN judd jr., a.f. BISH 10/1/1931
115 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2749 Kauai PTBG 2/5/1988
116 Ophioglossum polyphyllum unknown Lowland Dry Shrubland at 1800 ft on a W facing slope with Dodonaea viscosa, Sida fallax, Lipochaeta lavarum, L. lobata and Eragrostis sp. PRESERVED_SPECIMEN munro, g.c. collector number: 167 Maui BISH
117 Ophioglossum polyphyllum Kamoku. PRESERVED_SPECIMEN Munro, G.C. 167 Lanai BISH 4/2/1905
118 Ophioglossum polyphyllum Wailuku PRESERVED_SPECIMEN Bailey, E. s.n. Maui BISH 1879-00-00
119 Ophioglossum polyphyllum PRESERVED_SPECIMEN d. d. baldwin 113 MU
120 Ophioglossum polyphyllum Polihale State Park, dunes in front of reef zone near Queens Pond. Coastal dry shrubland mixed community, sand dunes and lithified sand outcrops. PRESERVED_SPECIMEN K. R. Wood 18381 Kauai PTBG 2/11/2020
121 Ophioglossum polyphyllum N. Kona; upper makai slopes of Puu Oo, Land of Maniniowale, mauka of Queen Kaahumanu Hwy Ca. 100 individuals near summit, in exposed sites in grassland of Pennisetum setacum. PRESERVED_SPECIMEN Nagata, K.M. 4270 Hawaii BISH 11/22/1993
122 Ophioglossum polyphyllum unknown PRESERVED_SPECIMEN Judd, J.R. s.n. BISH 1888-00-00
123 Ophioglossum polyphyllum n. kona; upper makai slopes of puu oo, land of maniniowale, mauka of queen kaahumanu hwy PRESERVED_SPECIMEN nagata, k.m. collector number: 4270 Kauai BISH 11/22/1993
124 Ophioglossum polyphyllum Kanaio, near Piimoe Hill, above Pohakea; slopes all the way to Hwy have scattered plants, eastern side of Kanaio. Elev. 1170-1460 ft. Alien vegetation. PRESERVED_SPECIMEN Steve Perlman 13309 Maui PTBG 2/4/1993
125 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2683 Kauai PTBG 1/15/1988
126 Ophioglossum polyphyllum Polihale State Park. Dune system behind beach. Low shrubland. On open dunes or in sand pockets of lithified dunes. PRESERVED_SPECIMEN Tim Flynn 3285 Kauai PTBG 3/7/1989
127 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2698 Kauai PTBG 1/18/1988
128 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2697 Kauai PTBG 1/18/1988
129 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2706 Kauai PTBG 1/18/1988
130 Ophioglossum polyphyllum Polihale State Park, dunes in front of reef zone near Queens Pond. Coastal dry shrubland mixed community, sand dunes and lithified sand outcrops. PRESERVED_SPECIMEN K. R. Wood 15850 Kauai PTBG 2/27/2014
131 Ophioglossum polyphyllum U. S. Navy Pacific Missile Range Facility at Barking Sands. PRESERVED_SPECIMEN Tim Flynn 5185 Kauai PTBG 1/11/1993
132 Ophioglossum polyphyllum East Maui, Keokea, SE of Puu O Kali. Lava flow. PRESERVED_SPECIMEN Hank Oppenheimer H111320 Maui PTBG 11/25/2013
133 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2700 Kauai PTBG 1/18/1988
134 Ophioglossum polyphyllum US Navy Pacific Missle Range Facility. Barking Sands. Just north of Nohili Point; base of sand dunes 200 ft inland from beach. Open areas with Leucaena-Prosopis shrubland. PRESERVED_SPECIMEN Walter Odening s.n. Kauai PTBG 1/31/1990
135 Ophioglossum polyphyllum Kau, Hilea-Kai, radius of 10' on heiau ~30' from ocean, full sun, dry PRESERVED_SPECIMEN Greenwell, A.B.H. Degener 21241 Hawaii BISH 3/9/1950
136 Ophioglossum polyphyllum ditch in the center of the oahu country club dried ditch PRESERVED_SPECIMEN d.f. grether 4234 WIS 2/22/1946
137 Ophioglossum polyphyllum Kamakaipo sand dunes PRESERVED_SPECIMEN Cooke Jr., C.M. 1268a Molokai BISH 1/1/1924
138 Ophioglossum polyphyllum Ninole, Wailua, Kau gravelly spot on basalt point, by old Hawaiian house site PRESERVED_SPECIMEN St. John, H. 23952 Hawaii BISH 12/25/1949
139 Ophioglossum polyphyllum puako In partly shaded lawn area mowed weekly. PRESERVED_SPECIMEN funk, e.j. collector number: s.n. Oahu BISH 12/18/1992
140 Ophioglossum polyphyllum kamilo, ka`u PRESERVED_SPECIMEN Hawaii US
141 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2748a Kauai PTBG 2/5/1988
142 Ophioglossum polyphyllum Manele and Hulopoe Bays, between sand dunes, kiawe forest PRESERVED_SPECIMEN Stemmermann, R.L. 3715B Lanai BISH 2/18/1979
143 Ophioglossum polyphyllum barking sands, between kohole point and base housing Dodonaea shrubland on low, coastal sand dunes, in full sun or in thatch PRESERVED_SPECIMEN wilson, k.a. gustafson, r.; flynn, t. collector number: 1612 Kauai BISH 3/6/1989
144 Ophioglossum polyphyllum kahue, puna district In moist black soil, in sink hole 2m deep with Sida. Semi arid pahoehoe slopes. PRESERVED_SPECIMEN st.john, h. collector number: 25466 Hawaii BISH 12/24/1954
145 Ophioglossum polyphyllum manele and hulopoe bays, between sand dunes, kiawe forest PRESERVED_SPECIMEN stemmermann, r.l. herbst, d.r.; palmer, d.; corn, c. collector number: 3715b Lanai BISH 2/18/1979
146 Ophioglossum polyphyllum Kauai PRESERVED_SPECIMEN Brodie, A. s.n. Kauai BISH
147 Ophioglossum polyphyllum east maui, s coast, hanamanioa point fine coral and lava rubble PRESERVED_SPECIMEN hobdy, r.w. collector number: 2459 Maui BISH 12/1/1985
148 Ophioglossum polyphyllum Kau, Keauhou back of high tide mark PRESERVED_SPECIMEN Judd Jr., A.F. s.n. Hawaii BISH 10/1/1931
149 Ophioglossum polyphyllum barking sands, between kohole point and base housing Dodonaea shrubland on low, coastal sand dunes, under canopy PRESERVED_SPECIMEN flynn, t. lorence, d.h.; crum, e.; momohara, d. collector number: 2683 Kauai BISH 1/15/1988
150 Ophioglossum polyphyllum Barking Sands, between Kohole Point and base housing Dodonaea shrubland on low, coastal sand dunes, under canopy PRESERVED_SPECIMEN Flynn, T. 2683 Kauai BISH 1/15/1988
151 Ophioglossum polyphyllum W Maui, Wailuku Sand hills. PRESERVED_SPECIMEN s.n. Maui BISH
152 Ophioglossum polyphyllum near sisal PRESERVED_SPECIMEN cooke jr., c.m. collector number: 1763.o Oahu BISH 2/16/1912
153 Ophioglossum polyphyllum Kahue, Puna District In moist black soil, in sink hole 2m deep with Sida. Semi arid pahoehoe slopes. PRESERVED_SPECIMEN St. John, H. 25466 Hawaii BISH 12/24/1954
154 Ophioglossum polyphyllum behind cove W of Kanaio beach hot cinders PRESERVED_SPECIMEN Hobdy, R.W. 414 Maui BISH 12/27/1978
155 Ophioglossum polyphyllum hulopoe sand under Prosopis PRESERVED_SPECIMEN herbst, d.r. collector number: 6312 Lanai BISH 2/18/1979
156 Ophioglossum polyphyllum Kahue, Puna moist black soil, sink hole 2 m deep with Sida, semiarid pahoehoe slopes PRESERVED_SPECIMEN St. John, H. 25466 Hawaii BISH 12/24/1954
157 Ophioglossum polyphyllum behind cove w of kanaio beach hot cinders PRESERVED_SPECIMEN hobdy, r.w. collector number: 414 Maui BISH 12/27/1978
158 Ophioglossum polyphyllum ruins 2 miles S of Pohie Bay soil on pahoehoe PRESERVED_SPECIMEN Higashino, P.K. 10207 Hawaii BISH 3/26/1984
159 Ophioglossum polyphyllum near Sisal PRESERVED_SPECIMEN Cooke Jr., C.M. 1763.O Oahu BISH 2/16/1912
160 Ophioglossum polyphyllum oahu PRESERVED_SPECIMEN remy, m.j. collector number: 58 Oahu BISH 1851-01-01
161 Ophioglossum polyphyllum Hulopoe sand under Prosopis PRESERVED_SPECIMEN Herbst, D.R. 6312 Lanai BISH 2/18/1979
162 Ophioglossum polyphyllum unknown PreservedSpecimen Judd, J.R. Collector Number: s.n. BISH Specimen
163 Ophioglossum polyphyllum ninole, wailua, kau gravelly spot on basalt point, by old Hawaiian house site PRESERVED_SPECIMEN st.john, h. hatheway, w.h.; morton, n.e. collector number: 23952 Hawaii BISH 12/25/1949
164 Ophioglossum polyphyllum Puako Under buffelgrass [Cenchrus ciliaris L.] PRESERVED_SPECIMEN Funk, E.J. s.n. Hawaii BISH 12/18/1992
165 Ophioglossum polyphyllum East Maui, Wailuku District, Keokea, southeast of Puu O Kali. Vegetation: Erythrina, Diospyros, Sophora, Myoporum, Euphorbia, Sideroxylon, Polyscias, Nothocestrum, Sida, Chenopodium, Nototrichium, Dodonaea, Capparis, Cocculus, Sicyos, Pellaea, Doryopteris, Ipomoea, Argemone, Waltheria, Prosopis, Melia, Leucaena, Neonotonia, Melinis, Verbesina, Lantana. PRESERVED_SPECIMEN Oppenheimer, H.L. H111320 Maui BISH 11/25/2013
166 Ophioglossum polyphyllum w maui, wailuku Sand hills. PRESERVED_SPECIMEN bailey, e. collector number: s.n. Maui BISH
167 Ophioglossum polyphyllum unknown PreservedSpecimen Munro, G.C. Collector Number: 167 BISH Specimen 4/2/1905
168 Ophioglossum polyphyllum unknown PreservedSpecimen Kanashiro, T. Collector Number: 5730 BISH Specimen 12/11/1947
169 Ophioglossum polyphyllum kau, hilea-kai, radius of 10' on heiau ~30' from ocean, full sun, dry PRESERVED_SPECIMEN greenwell, a.b.h. collector number: degener 21241 Hawaii BISH 3/9/1950
170 Ophioglossum polyphyllum East Maui, S coast, Hanamanioa Point fine coral and lava rubble PRESERVED_SPECIMEN Hobdy, R.W. 2459 Maui BISH 12/1/1985
171 Ophioglossum polyphyllum ruins 2 miles s of pohie bay PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10207 BISH 3/26/1984
172 Ophioglossum polyphyllum kamakaipo sand dunes PRESERVED_SPECIMEN cooke jr., c.m. collector number: 1268a Molokai BISH 1/1/1924
173 Ophioglossum polyphyllum hawaiian islands. PRESERVED_SPECIMEN d. d. baldwin s.n. GH
174 Ophioglossum polyphyllum kauai Exposed slopes, mostly bare ground with sparse scattering of wind-trimmed Schinus, Metrosideros & Styphelia PRESERVED_SPECIMEN brodie, a. Oahu BISH
175 Ophioglossum polyphyllum Barking Sands, between Kohole Point and base housing Dodonaea shrubland on low, coastal sand dunes, in full sun or in thatch PRESERVED_SPECIMEN Wilson, K.A. 1612 Kauai BISH 3/6/1989
176 Ophioglossum polyphyllum w maui, wailuku sand hills PRESERVED_SPECIMEN Maui BISH
177 Ophioglossum polyphyllum south kohala district. collected east of queen kaahumanu highway between mauna lani drive and puako turn off Growing in shady forest on steep, talus slope near valley bottom. Associated vegetation: Aleurites moluccana, Psidium guajava, Pouteria sandwicensis, Freycinetia arborea, Antidesma platyphyllum, Pipturus albidus, Cyrtandra grayi, Cyrtandra munroi, Tectaria gaudichaudii, Strongylodon ruber, Christella parasitica, Blechnum appendiculatum. PRESERVED_SPECIMEN funk, e.j. collector number: s.n. Maui BISH 2/4/1991
178 Ophioglossum polyphyllum W Maui, Wailuku Sand hills. PRESERVED_SPECIMEN Bailey, E. s.n. Maui BISH
179 Ophioglossum polyphyllum unknown PRESERVED_SPECIMEN judd, j.r. collector number: s.n. Oahu BISH
180 Ophioglossum polyphyllum Oahu PRESERVED_SPECIMEN Remy, M.J. 58 Oahu BISH 1/24/1905
181 Ophioglossum polyphyllum kau, keauhou PRESERVED_SPECIMEN judd jr., a.f. BISH 10/1/1931
182 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2749 Kauai PTBG 2/5/1988
183 Ophioglossum polyphyllum unknown Lowland Dry Shrubland at 1800 ft on a W facing slope with Dodonaea viscosa, Sida fallax, Lipochaeta lavarum, L. lobata and Eragrostis sp. PRESERVED_SPECIMEN munro, g.c. collector number: 167 Maui BISH
184 Ophioglossum polyphyllum Kamoku. PRESERVED_SPECIMEN Munro, G.C. 167 Lanai BISH 4/2/1905
185 Ophioglossum polyphyllum Wailuku PRESERVED_SPECIMEN Bailey, E. s.n. Maui BISH 1879-00-00
186 Ophioglossum polyphyllum PRESERVED_SPECIMEN d. d. baldwin 113 MU
187 Ophioglossum polyphyllum Polihale State Park, dunes in front of reef zone near Queens Pond. Coastal dry shrubland mixed community, sand dunes and lithified sand outcrops. PRESERVED_SPECIMEN K. R. Wood 18381 Kauai PTBG 2/11/2020
188 Ophioglossum polyphyllum N. Kona; upper makai slopes of Puu Oo, Land of Maniniowale, mauka of Queen Kaahumanu Hwy Ca. 100 individuals near summit, in exposed sites in grassland of Pennisetum setacum. PRESERVED_SPECIMEN Nagata, K.M. 4270 Hawaii BISH 11/22/1993
189 Ophioglossum polyphyllum unknown PRESERVED_SPECIMEN Judd, J.R. s.n. BISH 1888-00-00
190 Ophioglossum polyphyllum n. kona; upper makai slopes of puu oo, land of maniniowale, mauka of queen kaahumanu hwy PRESERVED_SPECIMEN nagata, k.m. collector number: 4270 Kauai BISH 11/22/1993
191 Ophioglossum polyphyllum Kanaio, near Piimoe Hill, above Pohakea; slopes all the way to Hwy have scattered plants, eastern side of Kanaio. Elev. 1170-1460 ft. Alien vegetation. PRESERVED_SPECIMEN Steve Perlman 13309 Maui PTBG 2/4/1993
192 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2683 Kauai PTBG 1/15/1988
193 Ophioglossum polyphyllum Polihale State Park. Dune system behind beach. Low shrubland. On open dunes or in sand pockets of lithified dunes. PRESERVED_SPECIMEN Tim Flynn 3285 Kauai PTBG 3/7/1989
194 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2698 Kauai PTBG 1/18/1988
195 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2697 Kauai PTBG 1/18/1988
196 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2706 Kauai PTBG 1/18/1988
197 Ophioglossum polyphyllum Polihale State Park, dunes in front of reef zone near Queens Pond. Coastal dry shrubland mixed community, sand dunes and lithified sand outcrops. PRESERVED_SPECIMEN K. R. Wood 15850 Kauai PTBG 2/27/2014
198 Ophioglossum polyphyllum U. S. Navy Pacific Missile Range Facility at Barking Sands. PRESERVED_SPECIMEN Tim Flynn 5185 Kauai PTBG 1/11/1993
199 Ophioglossum polyphyllum East Maui, Keokea, SE of Puu O Kali. Lava flow. PRESERVED_SPECIMEN Hank Oppenheimer H111320 Maui PTBG 11/25/2013
200 Ophioglossum polyphyllum Barking Sands. Pacific Missle Range Facility, US Navy. Between Kokole Point and base housing. In Dodonaea shrubland on low, coastal sand dunes. PRESERVED_SPECIMEN Tim Flynn 2700 Kauai PTBG 1/18/1988
201 Ophioglossum polyphyllum US Navy Pacific Missle Range Facility. Barking Sands. Just north of Nohili Point; base of sand dunes 200 ft inland from beach. Open areas with Leucaena-Prosopis shrubland. PRESERVED_SPECIMEN Walter Odening s.n. Kauai PTBG 1/31/1990
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